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Species images and videos
Tourists and visitors that undertake activities at Semuliki are encouraged to take videos or images of butterflies that they come across which all helps to contribute to the knowledge of the butterfly fauna of the park. Any images or videos used within the website will be acknowledged as will any new or important information that contributes further to the butterfly diversity of the park.

Species record additions and amendments

A provisional total of 448 butterfly species were documented and recorded at Semuliki National Park (Forbes, 2018). The park has by far the greatest butterfly diversity of any park or reserve in Uganda. There have been further additional species identified since the 2018 publication (currently 605 species - 05/04/2024). This has been through the further analysis of the thousands of images collected from the PhD research; analysing the excellent individual genus publications of Mark Williams; the genus revisions of Michel Libert and the unpublished Hesperiidae document of Torben Larsen. Not forgetting the continuing sampling of Justice Oribokihiro in the field prior to his transfer to Kibale National Park. The following additional records have therefore been collated which are in addition to what has been documented in Forbes (2018). The current confirmed recorded total now comes to 605 species.

Family Nymphalidae

Subfamily Charaxinae Doherty, 1886

Tribe Charaxini Guenée, 1865

Genus Charaxes Ochsenheimer, 1816

Charaxes smaragdalis leopoldi Ghesquière,1933 from Charaxes smaragdalis caerulea 

Subfamily Limenitidinae Behr, 1864

Genus Bebearia Hemming, 1960

Bebearia cocalia ssp. katera van Someren, 1939

Genus Acraea Fabricius, 1807

Subfamily Heliconiinae Swainson, 1822

Acraea paragea Grose-Smith, 1900 (ref: Acraeinae of Uganda (Bernaud et al., 2018))  from A. parageum parageum.

The following newly recorded species below are referenced within the website and included in the updated species total:

Family Nymphalidae

Subfamily Satyrinae Boisduval, 1833

Tribe Satyrini Boisduval, 1833

Genus Bicyclus Kirby, 1871

Bicyclus mesogenina Grünberg, 1911 (Aduse-Poku et al., 2016)

Subfamily Limenitidinae Behr, 1864

Tribe Adoliadini Doubleday, 1845
Genus Euphaedra Hübner, 1819

Euphaedra intermedia Rebel, 1914 (Forbes capture) 
Euphaedra ueleana Hecq, 1982 (Forbes capture) 
Euphaedra ansorgei Hecq, 1974 (Forbes capture) 
Euphaedra simplex Hecq, 1978 (Forbes capture)
Euphaedra sinuosa sinuosa Hecq, 1974 (Forbes capture)

Euphaedra procera/subprocera Hecq, 1984 (Forbes capture)

Genus Bebearia Hemming, 1960

Bebearia chloeropsis Bethune-Baker, 1908 (Afrotropical Butterflies, Williams 2022)

Genus Euriphene Boisduval, 1847

Euriphene doriclea ducarmei, Hecq, 1994 (Hecq, 2002)

Tribe Cymothoini Dhungel & Wahlberg, 2018

Genus Cymothoe Hübner, 1819

Cymothoe distincta Overlaet, 1952 (Afrotropical Butterflies, Williams 2022) - included in updated species total

Tribe Neptini Newman, 1870
Neptis Fabricius, 1807

Neptis agouale parallela Collins & Larsen, 1996 is replaced by Neptis melicerta parallela (Collins & Larsen, 1996 comb. nov.)


Subfamily Heliconiinae Swainson, 1822
Tribe Acraeini Boisduval, 1833
Subtribe Acraeina Boisduval, 1833

Genus Acraea Fabricius, 1807

Acraea endoscota Le Doux, 1928 (ref: Acraeinae of Uganda (Bernaud et al., 2018))

Acraea ventura Hewitson, 1877 (ref: Aurivillius, 1925 - Acraeinae of Uganda (Bernaud et al., 2018))

Subfamily Biblydinae

Tribe Epicaliini Guenée, 1865

Genus Sevenia Koçak, 1996
Sevenia umbrina Karsch, 1892 (Forbes capture)


Family Hesperiidae

Subfamily Tagiadinae Mabille, 1878

Tribe Tagiadini Mabille, 1878

Genus Eagris Guenée, 1862

Eagris tetrastigma tetrastigma Mabille, 1891 (Forbes capture)

Eagris tigris tigris Evans, 1937

Eagris nottoana nottoana Wallengren,1857

Genus Abantis Hopffer, 1855

Abantis iruma Evans, 1951

Tribe Celaenorrhinini Swinhoe, 1912

Genus Sarangesa Moore, 1881

Sarangesa tertullianus Fabricius, 1793

Sarangesa bouvieri Mabille, 1877

Sarangesa maculata Mabille, 1891

Sarangesa thecla mabira Evans, 1956

Sarangesa loelius Mabille, 1877

Sarangesa haplopa Swinhoe, 1907

Genus Scopulifera Libert, 2014

Scopulifera nigropuncta Bethune-Baker, 1908

Genus Bettonula Libert & Larsen, 2014

Bettonula bettoni bettoni Butler, 1902

Genus Apallaga Strand, 1911

Apallaga fulgens montanus Libert, 2014

Apallaga homeyeri fracta Libert, 2014

Apallaga inexpectata  Libert, 2014

Apallaga darlustris Libert, 2014

Apallaga alluaudi Mabille & Boullet, 1916

Apallaga boadicea howarthi Berger, 1976

Genus Eretis Mabille, 1891

Eretis buamba Evans, 1937

Eretis melania Mabille, 1891

Eretis artorius Larsen & Cock, 2011

Eretis camerona Evans, 1937

Genus Celaenorrhinus Hübner, 1819

Celaenorrhinus proxima proxima Mabille, 1877 (Forbes capture)

Celaenorrhinus perimitans Libert, 2014

Celaenorrhinus macrostictus Holland, 1893

Genus Spialia Swinhoe, 1912

Spialia ploetzi ploetzi Aurivillius, 1891

Subfamily Hesperiinae Latreille, 1809

Tribe Hesperiini Latreille, 1809

Genus Lepella Evans, 1937

Lepella lepeletier  Latreille, 1824

Genus Prosopalpus Holland, 1896

Prosopalpus styla Evans, 1937

Genus Gorgyra Holland, 1896

Gorgyra diversata Evans, 1937

Gorgyra mocquerysii Holland, 1896

Genus Teniorhinus Holland, 1892

Teniorhinus watsoni Holland, 1892 (recorded in error)

Teniorhinus niger  Druce, 1910

Genus Xanthodisca Aurivillius, 1925

Xanthodisca vibius Hewitson, 1878

Xanthodisca  astrape Holland, 1892

Genus Hypoleucis Mabille 1891

Hypoleucis dacena Hewitson, 1876

Hypoleucis ophiusa ophir Evans, 1937

Genus Andronymus Holland, 1896

Andronymus gander Evans, 1947

Andronymus caesar philander Hopffer, 1855

Genus Zophopetes Mabille, 1904

Zophopetes cerymica Hewitson, 1867

Zophopetes ganda Evans, 1937

Genus Fresna  Evans, 1937

Fresna netopha Hewitson, 1878

Fresna nyassae Hewitson, 1878

Fresna joppa Evans, 1937

Tribe Baorini Doherty, 1886

Genus Zenonia Evans, 1935

Zenonia zena Trimen, 1864

Zenonia crasta Evans, 1937

Genus Pelopidas Walker, 1870

Pelopidas thrax Hübner, 1821

Pelopidas mathias mathias Fabricius, 1798

Genus Borbo Evans, 1949

Borbo lugens Hopffer, 1855

Borbo  detecta Trimen, 1893

Borbo  borbonica borbonica Boisduval, 1833

Borbo  fatuellus fatuellus Hopffer, 1855

Borbo micans Holland, 1896

Tribe Ceratrichiini Grishin, 2019

Genus Ceratrichia Butler, 1870

Ceratrichia hollandi Bethune-Baker, 1908

Ceratrichia enta Evans, 1947

Genus Pardaleodes Butler, 1870

Pardaleodes bule Holland, 1896

Genus Ankola Evans, 1937

Ankola fan Holland, 1894

Tribe Astictopterini Swinhoe, 1912

Genus Osmodes Holland, 1892

Osmodes banghaasi Holland, 1896

Osmodes adonia Evans, 1937

Osmodes adosus Mabille, 1889

Osmodes lux Holland, 1892

Osmodes hollandi Evans, 1937

Genus Caenides Holland, 1896

Caenides kangvensis Holland, 1896

Genus Monza Evans, 1937

Monza cretacea Snellen, 1872

Monza alberti Holland, 1896

Genus Rhabdomantis Aurivillius, 1925

Rhabdomantis galatia Hewitson, 1868

Genus Acleros Mabille, 1885

Acleros bibundica Strand, 1913

Acleros nigrapex Strand, 1913

Genus Semalea Holland, 1896

Semalea pulvina Plötz, 1879

Semalea sextilis Plötz, 1886

Genus Chondrolepis Mabille, 1904

Chondrolepis niveicornis Plötz, 1883

Genus Paronymus Aurivillius, 1925

Paronymus ligora Hewitson, 1876

Genus Mopala Evans, 1937

Mopala orma Plötz, 1879

Genus Pteroteinon Watson, 1893

Pteroteinon caenira Hewitson, 1867

Pteroteinon capronnieri  Plötz, 1879

Tribe Gretnini Grisham, 2019

Genus Gretna Evans, 1937

Gretna zaremba jacksoni Evans, 1937

Family Lycaenidae

Species record additions for the Family Lycaenidae have been referenced from the publications of individual genera  found in  Afrotropical Butterflies (Williams, 2022). 

Subfamily Poritiinae Doherty, 1886

Tribe Pentilini Aurivillius, 1914

Genus Telipna Aurivillius, 1895

Telipna acraea nigrita Jackson, 1969

Telipna citrimaculata neavei Bethune-Baker, 1926

Telipna erica Suffert, 1904

Genus Pentila Westwood, 1851

Pentila hiendlmayri Dewitz, 1887

Pentila hybrida Libert and Collins, 2019

Genus Ornipholidotos Bethune-Baker, 1914

Ornipholidotos ugandae ugandae Stempffer, 1947

Ornipholidotos ntebi Bethune-Baker, 1906

Ornipholidotos paradoxa orientis Libert, 2005

Tribe Liptenini Röber, 1892

Genus Liptena Westwood, 1851

Liptena flavicans katera Stempffer, 1956

Liptena batesana Bethune-Baker, 1926

Liptena decipiens leucostola Kirby, 1890

Genus Falcuna Stempffer & Bennett, 1963

Falcuna iturina Stempffer and Bennett, 1963

Genus Citrinophila Kirby, 1887

Citrinophila tenerea Kirby, 1887

Genus Mimacraea Butler, 1872

Mimacraea fulvaria fulvaria Aurivillius, 1895

Mimacraea telloides Schultze, 1923

Tribe Epitolini Jackson, 1962

Genus Iridana Aurivillius, 1920

Iridana bwamba Stempffer, 1964

Iridana obscura Stempffer, 1964

Iridana katera Stempffer, 1964

Genus Geritola Libert, 1999

Geritola liana Roche, 1954

Geritola dubia dubia Jackson, 1964

Geritola zelica viridis Libert, 2020

Genus Stempfferia Jackson, 1962

Stempfferia badura contrasta Libert, 1999

Stempfferia cercene Hewitson, 1873

Stempfferia ciconia mongiro Jackson, 1968

Stempfferia congoana orientalis Libert, 2020

Stempfferia insulana Aurivillius, 1923

Stempfferia sylviae Libert, 1999

Genus Cephetola Libert, 1999

Cephetola entebbeana Bethune-Baker, 1926

Cephetola cephenoides Libert, 2020

Cephetola maculata Hawker-Smith, 1926

Cephetola viridana Joicey and Talbot, 1921

Cephetola orientalis Roche, 1954

Cephetola kamengensis subgriseata Jackson, 1964

Cephetola mpangensis mpangensis Jackson, 1962

Cephetola dolorosa Roche, 1954

Genus Epitolina Aurivillius, 1895

Epitolina melissa Druce, 1888

Genus Hewitsonia Kirby, 1871

Hewitsonia kuehnei Collins and Larsen, 2008

Genus Powellana Bethune-Baker, 1908

Powellana cottoni  Bethune-Baker, 1908

Subfamily Aphnaeinae Distant, 1884

Genus Pseudaletis Druce, 1888

Pseudaletis antimachus Staudinger, 1887

Subfamily Polymmatinae Swainson, 1827

Tribe Lycaenesthini Toxopeus, 1929

Genus Anthene Doubleday, 1847

Anthene lunulata grosei Aurivillius, 1899

Anthene akoae albidior Libert, 2010

Anthene irumu Stempffer, 1948

Anthene eliasi Congdon, Kielland and Collins, 1998

Genus Neurellipes Bethune-Baker, 1910

Neurellipes makala Bethune-Baker, 1910

Neurellipes zenkeri zenkeri Karsch, 1895

Neurellipes lachares toroensis Stempffer, 1947

Neurellipes onias Hulstaert, 1924

Genus Triclema Karsch, 1893

Triclema obsoleta Stempffer, 1947

Triclema rufoplagata ituriensis Joicey and Talbot, 1921

Triclema lamias katerae d'Abrera, 1980

Tribe Polyommatini Swainson, 1827

Genus Thermoniphas Karsch, 1895

Thermoniphas. togara togara Plötz, 1880

Genus Oboronia Karsch, 1893

Oboronia ornata vestalis Aurivillius, 1895

Subfamily Theclinae Swainson, 1830

Tribe Iolaini Riley, 1958
Iolaus Hübner, 1819

Iolaus mongiro Stempffer, 1969

Iolaus creta Hewitson, 1878

Iolaus neavei katera Talbot, 1937

Tribe Deudorigini Doherty, 1886

Genus Deudorix Hewitson, 1863

Deudorix odana odana Druce, 1887

Genus Hypomyrina Druce, 1891

Hypomyrina nomenia Hewitson, 1874

Hypomyrina fournierae Gabriel, 1939

Genus Paradeudorix Libert, 2004

Paradeudorix canescens Joicey & Talbot, 1921

Genus Pilodeudorix Libert, 2004

Pilodeudorix congoana orientalis Stempffer, 1957

Pilodeudorix anetia Hulstaert, 1924

Pilodeudorix mera mera Hewitson, 1873

Pilodeudorix otraeda genuba Hewitson, 1875

Pilodeudorix dimitris indentata Libert, 2004

Pilodeudorix bwamba Stempffer, 1962

Pilodeudorix violetta Aurivillius, 1897

Pilodeudorix camerona katanga Clench, 1965

Family Pieridae

Subfamily Pierinae Swainson, 1820

Tribe Teracolini Reuter, 1896

Genus Colotis Hübner, 1819

Colotis aurigineus Butler, 1883 (Forbes capture)

Colotis danae eupompe Klug, 1829 (Forbes capture)

Colotis evarne Klug, 1829 (Forbes capture)

Species record confirmations and request
The following species were recorded from Semuliki in Dr Peter Howard's 1991 report 'Nature Conservation in Uganda's Tropical Forest Reserves' from reference collections and literature reviews and presumably were included in the 374 species from the 'Biodiversity Report for Semuliki' (1996) by Dr Tim Davenport and Dr Peter Howard. However, only 309 species were actually identified in the Biodiversity Report list and these 8 species are the only ones I can find of the missing 65 species. It would be great to get images of the following from a museum or in the field:

Papilio antimachus - found at Kibale and DRC Semliki Valley (Ducarme) so definitely possible
Papilio leucotaenia - found in Kibale but montane habitat seems to be a preference
Papilio jacksoni ruandana - found by Ducarme in DRC Semliki Valley so definitely possible
Papilio rex mimeticus - found by Ducarme in DRC Semliki Valley so possible again

Graphium ridleyanus - again from DRC Semliki Valley (Ducarme)

Graphium almansor uganda - again Ducarme, DRC Semliki Valley and also Kibale 

Graphium ucalegon - Central Forest Belt/Ituri (Ducarme) so again possible, and Kibale

Graphium latreillanus - Central Forest Belt/Ituri (Ducarme) so likely

Specific references:

Ackery, P.R.Smith, C.R. and Vane-Wright, R.I., (1995). Carcasson's African Butterflies: An annotated Catalogue of the Papilionoidea and Hesperioidea of the Afrotropical Region. British Museum (Natural History), London, U.K.

Bernaud, D., Lequeux, J.P. and Ziraye, M., (2019). Acraeinae of Uganda


Davenport, T.R.D. & Howard, P. (1996). Semliki National Park Biodiversity Report. Forest Dept., Kampala.

Davenport, P., (2003). Endemic butterflies of the Albertine Rift - an annotated checklist. Wildlife Conservation Society, Mbeya.

Ducarme, R., (2018). The butterflies (Lepidoptera: Papilionoidia) of the north-eastern Democratic Republic of Congo. Metamorphosis 29, 22-36.

Howard, P., (1991). Nature Conservation in Uganda's Tropical Forest Reserves. IUCN/WWF, Forestry Dept., Ministry of Environment Protection, Kampala.

Libert, M., (2014). Sur la taxonomie du genre Celaenorrhinus Hübner en Afrique (Lepidoptera, Hesperiidae). ABRI, Nairobi.

Williams, M.C., (2021). Classification of the Afrotropical Butterflies to Generic Level. Publication is available from Lepidopterists' Society of Africa website

The website of Dominique Bernaud provides an unrivalled resource on the genus Acraea

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